4612 was built by the GWR at Swindon Works in 1942.
BR motive power depot allocations since 1948.
| Date Arrived | Depot |
| January 1948 | St Philips Marsh |
| June 1948 | Swindon |
| August 1948 | Chippenham |
| September 1948 | Swindon |
| October 1948 | Chippenham |
| October 1948 | Swindon |
| December 1948 | Chippenham |
| June 1950 | Swindon |
| May 1961 | Taunton |
| December 1961 | St Philips Marsh |
| January 1962 | Swindon |
| August 1962 | St Philips Marsh |
| October 1962 | Neath |
| June 1965 | Cardiff East Dock |
It ended its days working in South Wales, before withdrawal in 1965 and being sent to Barry scrapyard having completed 427,707 miles whilst in service. It was bought by the Keighley and Worth Valley Railway (KWVR) in 1981 for spares for 5775. It (or what was left of it) was then bought privately in 1987 by Elaine and Ray Treadwell and extensively restored, latterly at the Flour Mill workshop in the Forest of Dean. After restoration was completed in 2001, 4612 was moved to the Bodmin and Wenford Railway, which had purchased it, and returned to service in GWR green livery in 2013 following a ten-year overhaul.
In July 2019 the locomotive returned to traffic after undergoing repairs to firebox cracks.
In August 2020 the locomotive moved on loan to the Nene Valley Railway.
| Home Base | Current Status | Owner |
| Bodmin and Wenford Railway | Operational | Bodmin & Wenford Railway Trust |
Preserved British Steam Locomotives 